Lab Focus

My research involves meshing field research with theoretical models to address critical questions in ecology and conservation biology. I believe that ecological theory will be strengthened if it is forced to help solve real-world problems, and that conservation biology involves difficult choices that demand quantitative approaches.

Research Areas


Selected Recent Publications *

  • Fagan, W.F., F. McBride, and L. Koralov. 2023. Reinforced diffusions as models of memory-mediated animal movement. Journal of the Royal Society: Interface.20, p.20220700.
  • Fagan, W.F., et al. 2022. Quantifying interdependencies in geyser eruptions at the Upper Geyser Basin, Yellowstone National Park. Journal of Geophysical Research – Solid Earth. 127, e2021JB023749.
  • Fagan, W.F., C. Saborio, T. Hoffman, E. Gurarie, R. S. Cantrell, and C. Cosner. 2022. What’s in a resource gradient ? Exploring alternative cues for foraging in dynamic environments via movement, perception, and memory. Theoretical Ecology. 15: 267-282.
  • Rappoport, D., A. Swain, W.F. Fagan, R. Dubayah, and D.C. Morton. 2022. Animal soundscapes reveal key markers of Amazon forest degradation from fire and logging. Proceedings of the National Academy of Sciences, USA. 119: e2102878119.
  • Zambrano, J., G. Arellano, N.G. Swenson, P.P.A. Staniczenko, J. Thompson, and W.F. Fagan. 2022. Analyses of three‐dimensional species associations reveal departures from neutrality in a tropical forest. Ecology. 103: e3681.
  • Swain, A., L. Fussell, and W.F. Fagan. 2022. Higher-order effects, continuous species interactions, and trait evolution shape microbial spatial dynamics. Proceedings of the National Academy of Sciences, USA. 119 (1).
  • Mainali, K., E. Slud, M. Singer, and W. F. Fagan. 2022. A better index for analysis of co-occurrence and similarity. Science Advances. 8(4), p.eabj9204.
  • Mallon, J., K. Bildstein, and W.F. Fagan. 2021. Inclement weather forces stopovers and prevents migratory progress for obligate soaring migrants. Movement Ecology. 9, no. 1 (2021): 1-9
  • Lewis, M.A., W.F. Fagan, M. Auger-Méthé, J. Frair, J.M. Fryxell, C. Gros, E. Gurarie, S. Healy, J.A. Merkle. 2021. Learning and animal movement. Frontiers in Ecology and Evolution. 9, 681704.
  • Lin, H.-Y., W.F. Fagan, and P.-E. Jabin. 2021. Memory-driven movement model for periodic migrations. Journal of Theoretical Biology. 508, p.110486.
  • Martinez-Garcia, R., C. H. Fleming, R. Seppelt, W. F. Fagan, and J. M. Calabrese. 2020. How range residency and long-range perception change encounter rates. Journal of Theoretical Biology. p.110267.
  • Barry, T., E. Gurarie, F. Cheraghi, I. Kojola, and W.F. Fagan. 2020. Does dispersal make the heart grow bolder? Variation in habitat selection across wolf life history. Animal Behavior, 166, pp.219-231.
  • Mainali, K., T. Hefley, L. Ries, and W.F. Fagan. 2020. Matching expert range maps with species distribution model predictions. Conservation Biology. 34(5), pp.1292-1304.
  • Thompson, P., W.F. Fagan, and P.P.A. Staniczenko. 2020. Predictor species: Improving assessments of rare species occurrence by modeling environmental co-responses. Ecology and Evolution. 10: 3293-3304.
  • Che-Castaldo, C., C. Crisafulli, J.G. Bishop, E.F. Zipkin, and W.F. Fagan. 2019. Disentangling herbivore impacts in primary succession by refocusing the plant stress and vigor hypotheses on phenology. Ecological Monographs. p.e01389.
  • Fagan, W. F., T. Hoffman, D. Dahiya, E. Gurarie, S. Cantrell, and C. Cosner. 2019. Improved foraging by switching between diffusion and advection: Benefits from movement that depends on spatial context. Theoretical Ecology. p 1-10.
  • Swain, A., and W.F. Fagan. 2019. Group size and decision-making: Experimental evidence for Minority Games in fish behavior. Animal Behavior. 155: 9-19.
  • Noonan, M.J., M.A. Tucker, C.H. Fleming, …52 other coauthors …, W.F. Fagan, T. Mueller, and J.M. Calabrese. 2019. A comprehensive analysis of autocorrelation and bias in home range estimation. Ecological Monographs. 89: p.e01344.
  • Bewick, S., P. Staniczenko, B. Li, D. Karig, and W.F. Fagan. 2017. Invasion speeds in microbial systems with toxin production and quorum sensing. Journal of Theoretical Biology. 420: 290-303.
  • Fagan, W. F., E. Gurarie, S. Bewick, A. Howard, S. Cantrell, and C. Cosner. 2017. Perceptual ranges, information gathering, and foraging success in dynamic landscapes. American Naturalist. 189: 474-489.
  • Zhou, J., and W.F. Fagan. 2017. A discrete time model for populations in habitats with time-varying sizes. Journal of Mathematical Biology. 1-56.
  • Bewick, S., F. Agusto, J. M. Calabrese, E. J. Muturi, and W.F. Fagan. 2016. Epidemiology of La Crosse virus encephalitis emergence, Appalachia Region, United States. Emerging Infectious Diseases. 22: 1921.
  • Teitelbaum, C., S. Converse, W.F. Fagan, K. Boehning-Gaese, R. O’Hara, A. Lacey, and T. Mueller. 2016. Experience drives innovation of new migration patterns of whooping cranes in response to global change. Nature Communications. 7:12793 DOI: 10.1038/ncomms12793
  • Bewick, S., R.S. Cantrell, C. Cosner, and W.F. Fagan. 2016. How resource phenology affects consumer population dynamics. American Naturalist, 187: 151-166.
  • Foss-Grant, A. P., E. F. Zipkin, J.T. Thorson, O. P. Jensen, and W.F. Fagan. 2016. Hierarchical analysis of phylogenetic variation in intraspecific competition across fish species. Ecology, 97:1724-1734.
  • Teitelbaum, C., W.F. Fagan, C.H. Fleming, G. Dressler, J.M. Calabrese, P. Leimgruber, and T. Mueller. 2015. How far to go? Determinants of migration distance of land mammals. Ecology Letters. 18: 545-552. (Cover Article)
  • Che-Castaldo, C., C. Crisafulli, J.M. Bishop, and W.F. Fagan. 2015. What causes female bias in the secondary sex-ratios of the dioecious woody shrub Salix sitchensis colonizing a primary successional landscape? American Journal of Botany. 102: 1309-1322.
  • Zeigler, S. and W.F. Fagan. 2014. Transient windows for connectivity in a changing world. Movement Ecology. 2: 1.
  • Martinson, H.M. and W.F. Fagan. 2014. How habitat fragmentation alters the intensity of trophic interactions: a meta-analysis of trophic disruption in arthropod food webs. Ecology Letters. 17: 1178-1189.
  • Fleming, C.H., J.M. Calabrese, T. Mueller, K.A. Olson, P. Leimgruber, and W.F. Fagan. 2014. From fine-scale foraging to home ranges: A semi-variance approach to identifying movement modes across spatiotemporal scales. American Naturalist. E154-E167.
  • Mueller,, T., R. O'Hara, R. Urbanek, S. Converse, and W. F. Fagan. 2013. Social learning and migratory performance. Science. doi/10.1126/science.1237139. (Cover Article)
  • Fagan, W.F., M. A. Lewis, M. Auger-Méthé, T. Avgar, S. Benhamou, G. Breed, L. LaDage, U. Schlägel, W. Tang, Y. Papastamatiou, J. Forester, and T. Mueller. 2013. Spatial memory and animal movement. Ecology Letters. DOI: 10.1111/ele.12165
  • Fagan, W.F., Y. Pearson, E. Larsen, J.B. Turner, H.J. Lynch, H.Staver, J. Turner, A. E. Noble, S. Bewick, and E. Goldberg. 2013. Phylogenetic prediction of the maximum per capita rate of population growth. Proceedings of the Royal Society, Series B. 280: 20130523
  • Berbert, J.M., and W.F. Fagan. 2012. How the interplay between individual spatial memory and landscape persistence can generate population distribution patterns. Ecological Complexity. 12: 1-12.
  • Lynch, H.J., R. Naveen, P.N. Trathan, W.F. Fagan. 2012. Spatially integrated assessment reveals widespread changes in penguin populations on the Antarctic Peninsula. Ecology 93:1367-1377.


For Dr. Bill Fagan's CV, go here.

For additional publications, go here.

For information on lab research areas, go here.

For the Timing of Mosquito Control application, go here


Lab News

  • New National Science Foundation - Navigating the New Arctic grant supports investigations of caribou demography and movement ecology (2021 - 2024)
  • New National Science Foundation – Mathematical Biology grant supports incorporating more biological reality in mathematical models of animal movement (2019-2022)
  • New National Science Foundation – Infrastructure Innovation for Biological Research grant supports development of a formal data integration framework that combines animal presence, presence-absence, and movement tracking data into a unified population distribution estimate (2019-2022)

From the gallery

Research crew at work Cave Millipede Gazelle Calf in Mongolia Lab members caving in WVA Willow Borer Hybrid Towhee