Variation in B&D: EV
Fecundity of adult spotted owls = 0.34
In a “normal” year: 34% of adult females have 1 female offspring.
In a “bad” year, EV results in decreased r: e.g., births = 34% - “x”
In a “good” year, EV results in increased r: e.g., births = 34% + “x”