BSCI 442                        Lecture 30:  I.  Review
                                                    II. Exercises
Fall 2006

I. SUMMARY/REVIEW

Four major stages in the life cycle of a plant

                                                SEEDLING
                                      A                                B
                          SEED                            VEGETATIVE PLANT
                                      D                                C

                                   REPRODUCTIVE PLANT

A. Seed germination
B. Development of the vegetative body
C. Development of the reproductive organs-flowers, pollination, fertilization
D. Embryogenesis to form a seed

ORDERLY DEVELOPMENT DEPENDS ON COORDINATION AND ARE SUBJECT TO CONTROL AT THREE LEVELS.
         1.  Programmed sequence of gene expression
         2.  A balance of hormones
         3.  Environmental signals

A.  SEED --> SEEDLING (Seed Germination)

 a)  BREAKING OF DORMANCY
          Temperature
          Water
          Light
          Hormones:  ABA Decr., GA increase. (e.g. BARLEY)

 b)  REQUIREMENTS FOR GROWTH
          Water
          Oxygen
          Temperature
          Mineral nutrients

Initially heterotrophic, mature seedling becomes photosynthetically-competent.
 

B.  SEEDLING ---> VEGETATIVE PLANT (Vegetative development)

 GROWTH & DIFFERENTIATION TO A PLANT WITH SPECIFIC ORGANS, TISSUES, CELL TYPES.   Growth & Differentiation DEPENDS ON the genetic program, environmental conditions (e.g. light) & intrinsic hormones.

 a)  Shoot apical meristem gives rise to leaves and stem.
LEAF development: into cell types with particular structure and function. e.g. mesophyll cell with chloroplasts

          STEM
  b) Root apical meristem gives rise to the root.

REQUIREMENTS FOR AUTOTROPHIC GROWTH
          Light
          Water
          Essential mineral nutrients
          Oxygen
          CO2

 UPTAKE AND TRANSPORT of Water and Nutrients

Ability to synthesize all the complex organic biomolecules using inorganic ions and simple compounds.
             Photosynthesis
             N assimilation

SYNTHESIS OF NEW ORGANIC COMPOUNDS NEEDED FOR GROWTH & DEV.
         CARBOHYDRATES : CELLULOSE, STARCH
         PROTEINS:  ENZYMES
         NUCLEIC ACIDS:  DNA, RNA
 

C.  MATURE VEGET. PLANT ---> REPROD. PLANT
(Reproductive Development)

 ENVIRONMENTAL STIMULI :
              LIGHT, TEMP, PHOTOPERIOD
             HORMONES
             NEW GENES ARE EXPRESSED

Transition from shoot apical meristem to Formation of the floral apical meristem 

FORMATION OF FLOWER: sepals, petals, stamen, carpel
          POLLINATION,
          FERTILIZATION

D.  REPRODUCTIVE PLANT --->  SEED (Embryogenesis)         

AFTER FERTILIZATION, SEED DEVELOPMENT BEGINS