Formulae for calculating H and Ne

37A

1 = proportion of H0 lost at each generation

2Ne

1 - 1 = proportion of H0 remaining after the first generation

2 Ne

Ht = H0 1 - 1 t = the absolute amount of H0 remaining after

2Ne t generations

Ne = 4 MF 1) unequal sex ratios or

M + F 2) nonrandom breeding

decrease Ne